Mona Lisa, with a different hand.
Curved strokes with bristled edges, angular palette-knife marks, and pointed petals. Each study uses 128 translucent shapes and the same starting seed and search settings. The reconstructions below are SVGs: zoom in to inspect the edges.

These are a small style comparison, using greedy fitting at 50% opacity. Studio also offers its existing optimization modes and mixes the new marks into Combo. Brush texture comes from the outline, so each mark remains vector geometry.